Cand. Scient (biochemistry) and Dr. Phil. (semiotics), b. 1942. Professor emeritus at Biological Institute, University of Copenhagen. Did work in experimental biochemistry in the 1970s. Research interests gradually turned towards questions of theoretical biology. From 1988 work has focused on the developing field of biosemiotics. Recent publictions include A Legacy of Living Systems: Gregory Bateson as Precursor to Biosemiotics, Springer 2008 (as editor), Biosemiotics. An Examination into the Signs of Life and the Life of Signs, University of Scranton Press 2008 and "Semiotic scaffolding: A biosemiotic link between sema and soma” in. K. Cabell and J. Valsiner (Eds.): The Catalyzing Mind.Annals of Theoretical Psychology 11, Springer 2014.